My last blog post concerned a 210-million-year-old, gap-filling dinosaur from Zimbabwe that is helping palaeontologists to reconstruct the early evolution and dispersal of dinosaurs.
This post concerns another gap-filling fossil, although one from much closer to the present—just 30 million years ago. It is a remarkably complete skeleton of the early canid Mesocyon coryphaeus, one of the most complete pre-Ice Age canid skeletons yet found in North America.
This specimen fills a different sort of gap. Mesocyon coryphaeus, first named in 1879, is relatively common in the John Day Formation of Oregon, but it was previously known principally from skulls and teeth. Its postcranial skeleton—the bones behind the skull—had remained almost entirely unknown. The new specimen therefore fills an important anatomical gap in our understanding of how early canids moved, hunted and responded to the changing environments of Oligocene North America.
The fossil itself has had a curious history. It was collected during the late 1980s from land administered by the Bureau of Land Management near what is now the John Day Fossil Beds National Monument. Encased in a protective plaster jacket, it entered the monument’s collection after initially being mistaken for an oreodont—an extinct North American even-toed ungulate sometimes loosely described as resembling a mixture of sheep, camel and pig.
During the 1990s, the skull was separated from the surrounding rock for display, whereupon its teeth and cranial anatomy revealed that it was actually a canid. Park palaeontologists began exposing the remainder of the skeleton in 2012, but preparation continued intermittently for years. By the time the fossil was fully uncovered in 2022, more than 500 hours of painstaking preparation had been devoted to it.
University of Michigan palaeontologist Anne E. Kort, together with Jennifer Cavin of John Day Fossil Beds National Monument and Xiaoming Wang of the Natural History Museum of Los Angeles County, formally described the specimen in a paper published online in the Journal of Paleontology on 29 May 2026.
Known as JODA 3366, the specimen includes a complete cranium, an almost complete vertebral column in front of the sacrum, all the major limb bones and parts of both the forefeet and hindfeet. Only the tail and portions of the hands and feet are missing. It even preserves part of the baculum, or penis bone, establishing beyond reasonable doubt that the animal was male.
































